A Primary Male Autosomal Linkage Map of the Horse Genome

Table 1.

Description of Markers Used in this Study

Locus Description Primer sequences Ho Phys. assign. Reference
Genes
Blood groups
EAA erythrocyte antigen A 50 Bailey et al. (1979)
EAD erythrocyte antigen D 37.5 Sandberg (1973)
EAQ erythrocyte antigen Q 37.5 Bowling et al. (1985)
Microsatellite
IGF2 insulin-like GGAGCACAGAACATGAAAAC 87.5 12q13 H. Ellegren (unpubl.)
 growth factor 2 AAATTTAATTGGCACAAACC Raudsepp et al. (1997)
PCR-RFLPs
MC1R melanocyte-stimulating hormone receptor 1 GATGGATCCTTCTGGGCTCCCTCAACTC 37.5 3p12 Marklund et al. (1996b)
GTAGTAGGCGATGAAGAGCGTGCT T. Raudsepp (unpubl.)
KIT mast cell growth factor receptor ATTTATTCCAACTTAGCGAACTGCAGC 25 3q21 Marklund et al. (1996b)
TCAGACATCTTCGTGGACAAGCAGAGG
Lear et al. (1997)
Proteins
A1BG α-1-B glycoprotein 37.5 Juneja et al. (1987)
ALB albumin 50 3q14.3 Andersson and Sandberg (1982) Godard et al. (1998)
CA carbonic anhydrase 37.5 Sandberg et al. (1968)
ES carboxylesterase 50 Andersson and Sandberg (1982)
HBA hemoglobin A 37.5 13pter Bowling et al. (1988) Oakenfull et al. (1993)
PGD 6-phosphogluconate 25 2p12–13 Andersson et al. (1984)
 dehydrogenase Gu et al. (1992)
PGM phosphoglucomutase 37.5 Dawson and Jaeger (1970)
TF transferrin 50 Gahne (1966)
RFLPs
FUCA α-L-1 fucosidase 37.5 this study
GLUT glucose transporter 12.5 this study
 type 1
LPL lipoprotein lipase 12.5 this study
MYL myosin light 75 this study
 polypeptide 1, 3
TYR tyrosinase 12.5 this study
SSCP
ELA–DRB equine leucocyte CTCTGCAGCACATTTCCTGGAG 25 20q14 Fraser and Bailey (1996)
 antigen class II CGCCGCTGCACCAGGAA
Ansari et al. (1988)
Anonymous loci
A-14 microsatellite CAGCTGGGTGACACAGAGAG 75 2q14–21 Marti et al. (1998)
GTCATCACTACTCCCTACAC
A-17 microsatellite GTGGAGAGATAAAAGAAGATCC 50 26q13–14 Marti et al. (1998)
GGCCACAAGGAATGAACACAC
ASB1 microsatellite AGCAGAAACCCACTCAAGCC 75 Breen et al. (1997)
GCATAATACCCTCAAGGTC
ASB2 microsatellite CCTTCCGTAGTTTAAGCTTCTG 87.5 15q21.3–23 Breen et al. (1997)
CACAACTGAGTTCTCTGATAGG
ASB3 microsatellite AATTCATCTCAGTGCTCTACCAGC 87.5 4q12–13 Breen et al. (1997)
TTCATTTTCTACATGCACTACAGC
ASB4 microsatellite TAAATTGTAAAAGCTGGAGCCG 100 9q16–18 Breen et al. (1997)
GCAAATAGTAGTTAAGTCCTC
ASB5 microsatellite TCGAGGAGCTCATGACCTGG 75 9q16–18 Breen et al. (1997)
TTGTACAACTCTCCACCATAGC
ASB6 microsatellite GGCACAGATGTTAGCTCAGC 75 10p13 Breen et al. (1997)
ATGGAACCAGCCTGGATTGC
ASB7 microsatellite CTGGAAATTACAGTGGTCTTCTGG 62.5 19q14–16 Breen et al. (1997)
AGGTTTTCAGGGGCTTGCGAAGC
ASB8 microsatellite GACAACGTGGCAGCTCACTGCC 37.5 1q16–17 Breen et al. (1997)
GCAAGTAAGCCATATGTGCATGCG
ASB9 microsatellite GTGCGCATGTATGTGCGTGCC 75 10q21–23 Breen et al. (1997)
ATTTCCACAAGGGACATGAGG
ASB10 microsatellite GTTGTCTAGGTGCAGAATCTGG 75 Breen et al. (1997)
GTTATGTCTCCCCTTTCTCTACC
ASB11 microsatellite CCACCTATGTGTTCAGTTCACC 50 19q21–22 Breen et al. (1997)
GCACCAATGTTTATAGACTCCC
ASB12 microsatellite TCAGCAATAGAAGCCAGCTCC 75 1q12–13 Breen et al. (1997)
TCCTATGGAGGTGACCTTCCC
ASB13 microsatellite CTCTGAAAGAGCAGGATTGG 37.5 2q14.3–21.2 Breen et al. (1997)
GTCTTCTAAGTGGTAAGAGCC
ASB14 microsatellite CTCCATGAATTCTCGCAGGTTGG 25 6q21 Breen et al. (1997)
CCATGGGCCATATGCACACTGC
ASB15 microsatellite GTCCCAAAGGGACTCAGGAAGG 37.5 15q21 Breen et al. (1997)
TGGATGCCAGTGCATAGACAG
ASB17 microsatellite GAGGGCGGTACCTTTGTACC 62.5 2p14–15 Breen et al. (1997)
ACCAGTCAGGATCTCCACCG
ASB18 microsatellite TGCAGACAAAGCTGGACACTC 50 Breen et al. (1997)
CTGCTGAGAAAGCTTCTGC
ASB19 microsatellite GAGTTGGAGCTCAAGTCTGTC 62.5 15q21.3–23 Breen et al. (1997)
GTTTAGCAACTACAGCGTAGG
ASB22 microsatellite GAGGAATGTGAAATACAGGAGG 75 4q21 Breen et al. (1997)
TTTGTGGTCTTCCGTGCACC
ASB23 microsatellite GAGGTTTGTAATTGGAATG 62.5 3q22 Irvin et al. (1998)
GAGAAGTCATTTTTAACACCT Lear et al. (1998)
ASB35 microsatellite ATGCATGAGCAGAGTGTTCTTCC 62.5 M. Breen et al.
TAGTACTTCTCTCTTAATATAAGC  (unpubl.)
ASB36 microsatellite GAACATGTAGTGTTTACTCTGCC 50 M. Breen et al.
GAAGGTTTGTTGGGTCTTACAAGG  (unpubl.)
ASB37 microsatellite CCTGCAACTTTTTCCCAGCC 12.5 13q11–12 Irvin et al. (1998)
GGCAGATGTTAGCTCATGGC Lear et al. (1998)
ASB38 microsatellite TGGGGTTGCCTTGGTTACC 62.5 Irvin et al. (1998)
TCAGAGGATGAGGCACAGC
B-8 microsatellite TCCTCAGTCCTTTCTCATGC 62.5 15q14–21 Marti et al. (1998)
AGCTGAAGGCAATCTGTACC
D-8 microsatellite TTTTTGTGTCTCAGGAGTGTG 62.5 11p12–13 Marti et al. (1998)
AGTCTGATGGTGGAGGAAGG
ECA-2 microsatellite TTCCCTCCCATGGTTATTTTTC 100 (1q2.1) Sakagami et al. (1995)
TCTCTACTTTCATATACATTTGG
ECA-3 microsatellite GGTTCACACAGGAGTCAGGGA 50 (2p1.3–4) Tozaki et al. (1995)
CCTTCTGGTTTGCCTCGTCTC
HLM3 microsatellite GAAGGTAGAAAAGGAGGGCTAGAAC 12.5 Vega-Pla et al. (1996)
TCTAGAGGACCATTCTCTGGGCTGTG
HMB1 microsatellite GTGTGTATGCTTCCCAACCCTT 50 Binns et al. (1995)
GTTATAAAGCACTATGATCTCA
HMB2 microsatellite GTGCCACCACCTCTGTGATT 50 Binns et al. (1995)
TGGAGAAGGATCTTGGGCTC
HMB3 microsatellite CAAACATCAGTTAAGAGTGA 75 Binns et al. (1995)
CTCTAATCCAGCAGTGTTCA
HMB4 microsatellite AACCGCCTGAGCAAGGAAGT 62.5 Binns et al. (1995)
GCTCCCAGAGAGTTTACCCT
HMB5 microsatellite ACGGACACATCCCTGCCTGC 75 Binns et al. (1995)
GCAGGCTAAGGAGGCTCAGC
HMB6 microsatellite GAAGATGTCCGCTTTGATAT 75 Binns et al. (1995)
CACTGGCACATCCAGATTTG
HMS1 microsatellite CATCACTCTTCATGTCTGCTTGG 62.5 Guérin et al. (1994)
TTGACATAAATGCTTATCCTATGGC
HMS2 microsatellite ACGGTGGCAACTGCCAAGGAAG 62.5 Guérin et al. (1994)
CTTGCAGTCGAATGTGTATTAAATG
HMS3 microsatellite CCAACTCTTTGTCACATAACAAGA 62.5 Guérin et al. (1994)
CAATCCTCACTTTTTCACTTTGTT
HMS5 microsatellite TAGTGTATCCGTCAGAGTTCAAAG 62.5 Guérin et al. (1994)
GCAAGGAAGTCAGACTCCTGGA
HMS6 microsatellite GAAGCTGCCAGTATTCAACCATTG 75 Guérin et al. (1994)
CTCCATCTTGTGAAGTGTAACTCA
HMS7 microsatellite CAGGAAACTCATGTTGATACCATC 75 Guérin et al. (1994)
TGTTGTTGAAACATACCTTGACTGT
HMS18 microsatellite CAACAATGAAAATTTGTCCTGTGC 62.5 Godard et al. (1997)
GTAAATGAGTAGACAATCATGAGG
HMS19 microsatellite CTAACCAGCACAGAATGAATGGC 25 4q21 Godard et al. (1997, 1998)
TAAAAGAACAGTGGAGAGTAAAGTG
HMS20 microsatellite TGGGAGAGGTACCTGAAATGTAC 100 Guérin and Bertaud (1996)
GTTGCTATAAAAAATTGTCTCCCTAC
HMS23 microsatellite GATCCAATATTGTAAACCCCGCC 37.5 Godard et al. (1997)
CCTTCATAACCCTTATTGCAGCC
HMS45 microsatellite TGTTACAGGTATTGGTAAACTGTGC 75 Godard et al. (1997)
GGAACAAGAAGAAATCACTAATGTC
HMS46 microsatellite GTCTCAGCCAAAAGGTATTCAAGC 50 Godard et al. (1997)
TGGCACCAATATAGGTCACCTGG
HMS47 microsatellite CCTGCTGAGGACCTTGGAAGCT 25 22q19 Godard et al. (1997, 1998)
ATGTATTTTCAAGTCTAATATCTGCC
HTG2 microsatellite GATTGGCAACAGATGTTAACTCGG 12.5 Ellegren et al. (1992)
CCCCATGAGAACTAACAATGTTAG
HTG3 microsatellite TAACCTGGGTGCAAAGCCACCCAT 62.5 Ellegren et al. (1992)
TCAGGGCCAATCTTCCTCAC
HTG4 microsatellite CTATCTCAGTCTTCATTGCAGGAC 100 Ellegren et al. (1992)
CTCCCTCCCTCCCTCTGTTCTC
HTG5 microsatellite TGCTAAGCCTCAGCACATACA 50 Ellegren et al. (1992)
TGGAAATAAGGTTAGCAGGGATGC
HTG6 microsatellite CCTGCTTGGAGGCTGTGATAAGAT 37.5 15q26–27 Ellegren et al. (1992)
GTTCACTGAATGTCAAATTCTGCT Godard et al. (1998)
HTG7 microsatellite CCTGAAGCAGAACATCCCTCCTTG 25 Marklund et al. (1994)
ATAAAGTGTCTGGGCAGAGCTGCT
HTG8 microsatellite CAGGCCGTAGATGACTACCAATGA 62.5 Marklund et al. (1994)
TTTTCAGAGTTAATTGGTATCACA
HTG9 microsatellite TGTGGGAAGAGTGTCAATAGCTGT 62.5 4q21.3 Marklund et al. (1994)
AGGCATCTGGTTTGCTGCAATTTC
Godard et al. (1998)
HTG10 microsatellite CAATTCCCGCCCCACCCCCGGCA 87.5 Marklund et al. (1994)
TTTTTATTCTGATCTGTCACATTT
HTG11 microsatellite CAATGATGGTACTTTGCATATTAA 37.5 Marklund et al. (1994)
ATCGGCATGCACACTCATAGGTAG
HTG12 microsatellite CACTAGAGTCAGGGGGGGTGGGCT 12.5 Marklund et al. (1994)
TTGGAGTACTCTTTCTCCCTTCCC
HTG13 microsatellite TTAGCACGGGGAGATCGGATCCTG 75 Marklund et al. (1994)
GGTCTCCCTCTCCATTCACCCTGC
HTG14 microsatellite CCAGTCTAAGTTTGTTGGCTAGAA 62.5 Marklund et al. (1994)
CAAAGGTGAGTGATGGATGGAAGC
HTG15 microsatellite TCTTGATGGCAGAGCCAGGATTTG 37.5 Marklund et al. (1994)
AATGTCACCATGCGGCACATGACT
HTG17 microsatellite GCTATCCCTCCTGAGTCTTA 87.5 Lindgren et al. (1998)
AGGTAATTTGAAATAAAATACAC
I-12 microsatellite AACTAAGCACGTCATACAAG 37.5 19q12–14 Marti et al. (1998)
CTTGTAGTTTTCGTTGTATAGC
I-18 microsatellite CAACAAAGATGTTGCAAGGG 50 16q23–25 Marti et al. (1998)
TGTGCCTCTTGTCTCTTAGG
LEX2 microsatellite AAAAGGAAGACTGGCGACAG 12.5 Coogle et al. (1996a)
GGTGGGGGAAAGAATGGT
LEX4 microsatellite AATAGCAAATCTCCCACTTCA 25 Coogle et al. (1996a)
GTCCTCACAACCTCATCATAA
LEX5 microsatellite AAGGCAATGCTTATCAAATGC 50 Coogle et al. (1996a)
TTACCCGCAGTGACTTCTATT
LEX7 microsatellite GGTAGGGCTCTGGGATGA 50 Coogle et al. (1996a)
AACACTGGGGAAAAGTCAG
LEX8 microsatellite AAACTGTCACAACGGTTAGGAC 37.5 Coogle et al. (1996a)
CGAAAAAGCCACTTGAGGTC
LEX9 microsatellite AAAGCCGTAAGATTGGGACA 75 Coogle et al. (1996a)
TCCATTGTGAGGGTGTAACA
LEX11 microsatellite ATTCCCAGTGAAGTATTGCCA 37.5 Coogle et al. (1996a)
AGAGATGGGTACCTGGGATTC
LEX14 microsatellite CCTTACTCACTGGGGAATAAA 87.5 Coogle et al. (1996a)
AGACTGAACACCTAACTATGA
LEX15 microsatellite GCATTCCCATCATCACAT 37.5 Coogle et al. (1996b)
CCTGCCTTGCCTCTTTCT
LEX16 microsatellite GTGGGGCCGGTATAGTGATTG 50 Coogle et al. (1996b)
ACCCTAACTGATAACTGATAGA
LEX17 microsatellite CCTGCCCAAGAAGAACTCAGA 100 Coogle et al. (1996b)
AGCAGTGTATTTTTGAAACAT
LEX18 microsatellite TTTCATCACTTTCTGCTTCC 25 Coogle et al. (1996b)
TTCTCTTCCTTTGCTCATCCT
LEX19 microsatellite TTCCCTTTTCCTCACATCCT 87.5 Coogle et al. (1996b)
TTTTAGGTTCATCTATGTTGTTGC
LEX20 microsatellite GGAATAGGTGGGGGTCTGTT 37.5 Coogle et al. (1996b)
AGGGTACTAGCCAAGTGACTGC
LEX21 microsatellite GTAGGCTTTCTGCCAAAAT 62.5 Coogle et al. (1996b)
TGAGGGGAGTCATAAAAA
LEX22 microsatellite AACATATCCATCGCCTCACA 25 Coogle et al. (1996b)
TGCAAATTCACTGAGAGTGG
LEX23 microsatellite GGATGAAACAGGGAAGGAAA 62.5 Coogle et al. (1996b)
CCAACGGATTCATGAAAGCTA
LEX25 microsatellite CAATCGTGGCCCGGTAAC 50 Coogle et al. (1996c)
TTCACTCCAATCCTCAGTCA
LEX27 microsatellite ACCACTGGGAAACTGTGTAA 25 Coogle et al. (1996c)
GCCCAGAATCCGAACC
LEX29 microsatellite TGGGGTGTCACTGCTTCTC 62.5 Coogle et al. (1996c)
ACTGAGGGCCAGGTTTCTAA
LEX30 microsatellite GGAGGGTGCAAGGTGCTA 37.5 Coogle et al. (1996c)
GGCAGGTCAGAAGGGACA
LEX31 microsatellite CCCATTAAGAACTTTTCATCCTG 87.5 Coogle et al. (1996c)
GGCAAGCCCCACAAAATTAT
LEX32 microsatellite CGTAGTAGGGTTTTGGGTCC 50 Coogle et al. (1996c)
TTGCGTTTCAATTTTTAATGAC
LEX33 microsatellite TTTAATCAAAGGATTCAGTTG 75 Coogle et al. (1996c)
TTTCTCTTCAGGTGTCCTC
LEX34 microsatellite GCGGAGGTAAGAAGTGGTAG 50 Coogle et al. (1997)
GGCCTAAGATGAGGGTGAA
LEX35 microsatellite CCCAGCATATCAAAGATGTT 75 Coogle et al. (1997)
GCTCAGTGTACTTCAAGCAG
LEX37 microsatellite GGATTCCTCAACCTCCTAAA 25 Coogle et al. (1997)
AGGGATAAGTGACCACCAC
LEX38 microsatellite CTGCATTCCCATCATCACAT 37.5 Coogle et al. (1997)
TGCCTTGCCTCTTTCTGTTTA
LEX39 microsatellite CCTCTGTCCCCACTACTCTC 37.5 Coogle et al. (1997)
TTGATCTCCACTCCCAATG
LEX40 microsatellite TTTGGCCGTTAGTCGTGT 37.5 Coogle et al. (1997)
GACAAATCGGAAAGTTGGAA
MPZ002 microsatellite GATCCCCCCTATTTTATATACAG 25 Breen et al. (1994)
AGGTTCTCATTCTACCTACAAGG
SGCV1 microsatellite AGTCACCACCACTCACCTTGT 37.5 ? Godard et al. (1997)
CCAACACAGGATACGGATGA
SGCV3 microsatellite CCTTGTGGTGAGTTTTCCTCTT 37.5 ? Godard et al. (1997)
CTGCAAAGCTCTGAAGGTC
SGCV4 microsatellite CGACGCCTCCTCCTAAAC 37.5 23q19 Godard et al. (1997)
CAGCTGTGTGCCTTTGATTAT
SGCV6 microsatellite GGGCCTGGTTTTCCTTCTAA 62.5 15q24 Godard et al. (1997)
GCATTTGTGGCCTGTGTCATA
SGCV7 microsatellite GAATTTGAATGTATCTATTCTGAATG 50 18q21 Godard et al. (1997)
GTGAGTTTTCAAGCTGGCATATTC
SGCV8 microsatellite GAGTTCATTCTTTTTCGTGGCTG 37.5 19 Godard et al. (1997)
GGAAACACCCTAAGTGTCCCTTG
SGCV10 microsatellite CATCCATCCTTTCCAGCTCGATATTC 62.5 12p13 Godard et al. (1997)
CAAGACCGTAACTCAGGAGCCC
SGCV13 microsatellite GGACTAAAGCCCAACCATCCAGC 75 11q12 Godard et al. (1997)
CTCACCAGTAAGGGGTTATGGGGC
SGCV14 microsatellite CCCCAGTGGTTCCATTTAGATGT 75 21q13 Godard et al. (1997)
GGGGAGAGCATTTTGGTGA
SGCV16 microsatellite AATTCTCAAATGGTTCAGTGA 37.5 21q13 Godard et al. (1997)
CTCCCTCCCTTCCTTCTA
SGCV17 microsatellite GGCCCAACGTCTATAGAAAGATGT 37.5 ? Godard et al. (1997)
CCCCCAAATGGCTATTTTCTAA
SGCV18 microsatellite TGGGGAAGAGGGATTCAT 62.5 3p13-14 Godard et al. (1997)
AAATGCCAAGCCTATCTATGC
SGCV19 microsatellite GCCCCCACCTGCTCCACC 62.5 22q19 Godard et al. (1997)
GGGGCAAAGTGGAAATCC
SGCV23 microsatellite GGCTTAAGATATGGGTGAGTAAGG 87.5 4q27 Godard et al. (1997)
GCCCACCCTCTTACTTTTCTCAA
SGCV24 microsatellite CTACCATTGAAGAGGGGTGGC 50 11p12 Godard et al. (1997)
GAAACGAGCAGGAAGTGAATCTCC
SGCV25 microsatellite GCCCATATTAGTAGGACTGTG 37.5 1q14 Godard et al. (1997)
GGCCATATTCAGCAGAGCT
SGCV28 microsatellite CTGTGGCAGCTGTCATCTTGG 25 Godard et al. (1997)
CCCAATTCCAGCCCAGCTTGC
SGCV30 microsatellite ACTGGAGGGGTGAAACAGATTCAGA 75 10qter Godard et al. (1997, 1998)
GGAAGGGAGGTCATCAGAA
SGCV32 microsatellite TGTTCCAAAATGGAGGGTGAGCC 62.5 ? Godard et al. (1997)
CCACAGGCTCTTAAAACCAGAAGC
VHL20 microsatellite CAAGTCCTCTTACTTGAAGACTAG 87.5 van Haeringen et al. (1994)
AACTCAGGGAGAATCTTCCTCAG
VIAS-H34 microsatellite GTATCAGCTTAACAGCTTTCTTTAAATG 37.5 Ewen and Matthews (1994a)
CTCCCGTCTCCTCTCTTGTTC
VIAS-H39 microsatellite AATGTGATTATAGCAGATAGGGTT 37.5 Ewen and Matthews (1994b)
CTATCCAATCTTCACAATCATGTA
  • Observed heterozygosity among the eight sires in the reference pedigree.

  • The assignment provided here is based on new experiments, which indicates that these clones are chimeric (M. Breen, unpubl.). Within the chimeric clones, the largest pieces of DNA map to the locations indicated in Breen et al. (1997). However, the sequence surrounding the microsatellite loci have now been confirmed to map to the locations given here.

  • Chromosome band identification was done using an old idiogram nomenclature.

  • Physical assignment does not fit with linkage data.

This Article

  1. Genome Res. 8: 951-966

Preprint Server