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A second calender year male flycatcher just arrived from his winter visit in Angola, Africa to the Swedish island Gotland—a trip of around 8500 km. In this issue, the germline mutation rate is estimated following deep resequencing of 11 members of a three-generation flycatcher pedigree. (Cover photo by Johan Träff. [For details, see Smeds et al., pp. 1211–1218.])

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