Biological attributes of different categories of overlapping genes
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All genes have been placed in homologous groups, and for each category we present the mean PDI, the number that are nucleocapsid or replicase genes, and (for overlapping genes) the number that are +1 and −1 frameshifted. For the PDI and proportion of nucleocapsid or replicase genes, the statistical significance of the difference between each value and that of non-overlapping genes is given (Wilcoxon sum of ranks test and contingency χ2 test, respectively); for the frameshift, the significance of the deviation from equality is given (goodness-of-fit χ2 test). Cutoff for homology is a BLAST E-value of 10−3. NA, not applicable; NS, not significant.
*P = 0.01–0.05.
**P = <0.001.











