
Axes of variation of principal components 1 through 4. PC1 indicates size; PC2 is a trade-off between large pelvis, thin distal limbs, and short feet (metatarsal and metacarpal) vs. smaller pelvis, thicker distal limbs, and longer feet; PC3 is a trade-off between long thin bones vs. short thick bones; PC4, pelvic shape, is a trade-off between a longer narrower pelvis vs. shorter wider pelvis (see radiographs in Fig. 2). PC2 skeletal traits relevant to Figure 5 are circled.











