Table 1.

Experimentally determined cis-regulatory modules in the HBB gene cluster.


Chr11 Start

Chr11 Stop

CRM name

Function in regulation

Conserved in mousea

Conserved in rata

Conserved in chickena

References
5276843 5277003 HS5 LCR, enhancer blocker Yes Yes No (Dhar et al. 1990; Yu et al. 1994; Bender et al. 1998; Li et al. 2002; Tanimoto et al. 2003; Wai et al. 2003)
5273728 5274016 HS4 LCR, modulate enhancer Yes Yes No (Pruzina et al. 1991; Fraser et al. 1993; Stamatoyannopoulos et al. 1995; Molete et al. 2001)
5271123 5271701 HS3.2 LCR, modulate enhancer Yes Yes No (Slightom et al. 1997; Molete et al. 2002)
5270665 5270727 HS3.1 LCR Yes Yes No (Philipsen et al. 1990; Shelton et al. 1997)
5270191 5270478 HS3 LCR, modulate enhancer Yes Yes No (Fraser et al. 1990, 1993; Philipsen et al. 1990, 1993; Jackson et al. 1996)
5266399 5266483 HS2_neg LCR, negative Yes Yes No (Cavallesco and Tuan 1997; Elnitski et al. 2001)
5266104 5266398 HS2_pos LCR, enhancer Yes Yes No (Ryan et al. 1989; Tuan et al. 1989; Fraser et al. 1990, 1993; Talbot et al. 1990)
5261203 5261826 HS1 LCR Yes Yes No (Tuan et al. 1985; Forrester et al. 1986; Fraser et al. 1990, 1993)
5258391 5258617 HBE1_NRA Negative No No No (Li et al. 1998)
5258291 5258390 HBE1_PRA Positive Yes No No (Li et al. 1998)
5257195 5257237 HBE1_NRB Negative Yes Yes No (Li et al. 1998)
5256999 5257195 HBE1_PRB Positive Yes Yes No (Li et al. 1998)
5255653 5255919 HBE1_up Negative Yes Yes No (Cao et al. 1989; Trepicchio et al. 1993; Watt et al. 1993)
5255484 5255652 HBE1_prom Promoter Yes Yes Yes (Allan et al. 1983; Gong et al. 1991; Yu et al. 1991)
5240524 5241054 HBG2_up Positive, negative Yes Yes No (Perez-Stable and Costantini 1990; Stamatoyannopoulos et al. 1993)
5240320 5240523 HBG2_prom Promoter Yes Yes Yes (Gimble et al. 1988; McDonagh et al. 1991; Stamatoyannopoulos et al. 1993)
5235600 5236122 HBG1_up Positive, negative Yes Yes No (Perez-Stable and Costantini 1990; Stamatoyannopoulos et al. 1993)
5235395 5235599 HBG1_prom Promoter Yes Yes Yes (Gimble et al. 1988; McDonagh et al. 1991; Stamatoyannopoulos et al. 1993)
5232674 5233423 HBG1_3′enh Enhancer No No No (Bodine and Ley 1987; Lloyd et al. 1994)
5220022 5220469 HBD_prom Promoter Yes Yes No (Tang et al. 1997)
5212610 5212865 HBB-prom Promoter Yes Yes Yes (Chao et al. 1983; Wright et al. 1984; Myers et al. 1986; Antoniou et al. 1988; Cowie and Myers 1988)
5210185 5210449 HBB_3′enh Enhancer Yes Yes No (Behringer et al. 1987; Trudel and Costantini 1987; Antoniou et al. 1988)
5190322
5190802
3′HS1
Enhancer blocker
Nob
Nob
No
(Fleenor and Kaufman 1993; Farrell et al. 2002; Bulger et al. 2003)

Coordinates are for hg16 (July 2003 freeze) of the human genome.

a Conserved in mouse, rat, or chicken means that this segment of human chromosome 11 aligns in a level 1 or level 2 chain in the nets of BLASTZ alignments with human sequences (Kent et al. 2003; Schwartz et al. 2003), which is an indication of aligning to an orthologous region. Alignments to nonorthologous regions are not included.

b About 30 bp aligns with a nonorthologous region of mouse chromosome 7.