Relationship between actinorhizal plants and groups of Frankia strainsa

aCompiled after Benson et al. (2004) and Swensen (1996).
bAccording to the classification of the Angiosperm Phylogeny Group (APG 1998); all of these orders fall in the “Eurosid I” group of eudicots.
cThe number of genera within the family is listed along with the number of genera nodulated. Not all genera within a family are capable of nodulation.
dThe Frankia cluster refers to the clusters of Frankia strains in Figure 1 typically found in root nodules of each plant genus. 1, alder and casuarina strains; 3, elaeagnus strains; 2, rosaceous strains; ND, not determined; (3) indicates rare detection of an elaeagnus strain in a nodule or in surface layers of a nodule (Benson et al. 2004).
(NA) North America; (SA) South America.